Preprints
Filtering by Subject: Physical Sciences and Mathematics
When Policy Shapes Selection: Anticipating Evolutionary Feedbacks in Conservation
Published: 2026-05-02
Subjects: Physical Sciences and Mathematics
Applied ecology and conservation increasingly operate in systems where ecological and evolutionary processes are tightly coupled and can unfold on management-relevant timescales, yet most policy still treats populations as evolutionarily static. As a result, interventions often generate unintended outcomes, from resistance evolution and harvest-induced trait shifts to hidden losses of adaptive [...]
Lead and slant on the geometry of coiling in gastropods
Published: 2026-05-01
Subjects: Ecology and Evolutionary Biology, Evolution, Life Sciences, Other Ecology and Evolutionary Biology, Physical Sciences and Mathematics
Molluscan shells have been studied with various geometric models. Here I show that lead angle, the defining slope of a conical helix, emerges as a more useful parameter in morphometric analyses and (adaptationist) interpretation of covariation in coiling parameters. The widely used apical semiangle becomes redundant and uninformative, a passive consequence of taxon-specific lead angles and [...]
Heterogeneity in Statistics: A Conceptual and Methodological Review
Published: 2026-04-15
Subjects: Physical Sciences and Mathematics
Heterogeneity—the presence of meaningful variation across observations, in models, and in inferences—is a foundational concept in statistics that has many meanings. This review synthesizes the evolution of the meanings, methodologies, and interpretations of the four dominant and interconnected types of heterogeneity: (1) heteroscedasticity (non-constant variance), historically treated as a [...]
The lens of the Sonic Holobiont. A perspective on acoustic influence on microbial communities and its application as an additional layer to the holobiont concept.
Published: 2026-03-12
Subjects: Arts and Humanities, Biochemistry, Biophysics, and Structural Biology, Biodiversity, Ecology and Evolutionary Biology, Life Sciences, Microbiology, Other Arts and Humanities, Other Physical Sciences and Mathematics, Physical Sciences and Mathematics
When studying micro and macro biomes in the quest for a more general understanding, we can hardly escape from a holistic perspective. At first, symbiosis was demonstrated to be a ubiquitous phenomenon in living cells, shaping evolutionary patterns across species at very different scales. The “holobiont” concept gains a central role in modern biology. The observation of the complex inter- and [...]
Addressing Unobserved Covariates in Species Distribution Models: Inferential Impacts and Mitigation via Joint Species Distribution Models
Published: 2026-03-05
Subjects: Life Sciences, Physical Sciences and Mathematics
Species distribution models (SDMs) are widely used in ecology to assess the distribution of species populations across space and time. Correlative SDMs, in particular, are used to infer relationships between species records and environmental variables. A classical approach for implementing this type of SDMs is to employ generalized linear mixed models (GLMMs) as a parametric regression method. [...]
The Global Lakes Explorer: A basin-to-global scale data visualisation application for the assessment of nutrient emissions to lakes
Published: 2026-02-25
Subjects: Computer Sciences, Earth Sciences, Environmental Sciences, Life Sciences, Physical Sciences and Mathematics
Phosphorus and nitrogen are critical plant nutrients, essential for fertiliser production and global food security. However, poor management across the anthropogenic nutrient cycles leads to losses associated with pollution of water bodies, driving eutrophication, biodiversity loss, and methane emissions. Addressing these interconnected challenges requires a sustainable, integral, and where [...]
Biodiversity footprint of public procurement in Finland
Published: 2026-02-07
Subjects: Biodiversity, Environmental Indicators and Impact Assessment, Environmental Sciences, Life Sciences, Physical Sciences and Mathematics
Halting biodiversity loss requires systematic action from all sectors of society. The public sector is a significant actor globally in creating markets for goods and services, public procurement representing on average 13 – 20% of national gross domestic product. In this study, we assessed the biodiversity footprint of public procurement in Finland for years 2021 and 2022. We applied the [...]
Diversity in a fuzzy world: a review of models and measures
Published: 2026-02-06
Subjects: Life Sciences, Physical Sciences and Mathematics
Biological diversity metrics have evolved from simple species counts to measures that incorporate increasingly complex functional information. A critical aspect is that different families of diversity indices are grounded in distinct mathematical models for representing uncertainty, including classical set theory, probability theory, and fuzzy set theory, which shape what these measures can [...]
How does the rate of environmental change affect density-dependent population dynamics?
Published: 2026-01-20
Subjects: Dynamic Systems, Ecology and Evolutionary Biology, Life Sciences, Numerical Analysis and Computation, Physical Sciences and Mathematics, Population Biology
Natural populations experience variable environments. Anthropogenically driven environmental change, in particular, is expected to impose trends on key demographic parameters such as reproduction and survival. Theoretical studies of how such environmental changes affect populations have highlighted dynamical phenomena including bifurcation-related tipping points – typically identified by [...]
Dorsal Fin Edge Proportions and Their Relationship to Swimming Strategy in Sharks
Published: 2025-12-25
Subjects: Life Sciences, Physical Sciences and Mathematics
Dorsal fins play a critical role in elasmobranch locomotion by providing stability, reducing roll, and enhancing swimming efficiency. Variation in dorsal fin geometry may therefore reflect ecological niche and swimming strategy. This study quantitatively compares dorsal fin leading-to-lagging edge ratios in three shark species with differing ecological roles: the spiny dogfish (Squalus [...]
Reframing Population Genetic Structure as a Quantum Optimization Problem
Published: 2025-12-24
Subjects: Life Sciences, Physical Sciences and Mathematics
Population genetic structure is commonly inferred using statistical and ordination-based methods that emphasize variance partitioning or likelihood-based clustering. While powerful, these approaches may undersample the full space of possible population partitions, particularly in systems characterized by weak genetic differentiation and high connectivity. Here, I present a proof-of-concept [...]
Predicting unobserved driver of regime shifts in social-ecological systems with universal dynamic equations
Published: 2025-12-15
Subjects: Engineering, Physical Sciences and Mathematics, Social and Behavioral Sciences
Ecosystems around the world are anticipated to undergo regime shifts as temperatures rise and other climatic and anthropogenic perturbations erode the resilience of present-day states. Forecasting these nonlinear ecosystem dynamics can help stakeholders to prepare for the associated rapid changes. One major challenge is that regime shifts can be difficult to predict when they are driven by [...]
A Satellite-based Approach to Investigating Eutrophication in Lakes Receiving Wastewater Treatment Effluent: A Case Study of Lake Windermere
Published: 2025-12-11
Subjects: Life Sciences, Physical Sciences and Mathematics
Excessive phosphorus from the sewer systems in the UK is a major contributor to eutrophication in freshwater bodies. Chlorophyll-a levels are widely used as a reliable indicator of phosphorus driven eutrophication, and satellite remote sensing provides an effective means of monitoring these dynamics at high spatial and temporal resolution. This study evaluated the utility of Sentinel-2 imagery in [...]
A chronosequence of human remains on soil microbial populations
Published: 2025-12-09
Subjects: Life Sciences, Physical Sciences and Mathematics
Surficial human decomposition produces substantial and measurable shifts in soil chemistry and microbial composition. This decade-long investigation examined temporal changes beneath surface deposited human remains and identified strong microbial and chemical responses in the first twelve to twenty four months, including decreased microbial diversity, elevated soil nutrients and sustained [...]
Landscape drivers of population density of a vulnerable apex predator (Wilkerr/dingo)
Published: 2025-09-11
Subjects: Physical Sciences and Mathematics
Apex predators shape ecosystems globally, yet robust monitoring that assesses the effects of management actions and environmental variation on their populations is challenging. The dingo, Australia’s largest terrestrial predator, is ecologically and culturally significant. In many parts of Australia, dingoes now exist in fragmented and isolated populations, and our understanding of how their [...]