Alexa Fredston , Morgan Tingley, Montague Neate-Clegg, Luke J. Evans, Laura Antão, Natalie Ban, I-Ching Chen, Yi-Wen Chen, Lise Comte, David Edwards, Birgitta Evengard, Belen Fadrique, Sophie Falkeis, Robert Guralnick, David Klinges, Jonas J Lembrechts , Jonathan Lenoir , Juliano Palacios-Abrantes, Anibal Pauchard, Gretta Pecl, Malin Pinsky, Rebecca Senior, Jennifer Smith, Lydia Soifer, Jennifer Sunday, Ken Tape, Peter Washam, Brett Scheffers

Climate change is already leaving a broad footprint of impacts on biodiversity, from an individual caterpillar emerging earlier in spring to an entire plant community migrating poleward. Despite the various modes of how species are on the move, we primarily document shifting species along only one gradient (e.g., latitude or phenology) and along one dimension (space or time). Here we present a unifying framework for integrating the study of species on the move over space and time and from micro to macro scales. Future conservation planning and natural resource management will depend on our ability to use this framework to improve understanding, attribution, and prediction of species on the move.